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This is usually perceived as a taste on the sides of the tongue, towards the rear of the mouth. The two most common acids responsible for this flavor are lactic and acetic, which both have related esters that may be perceived in the aroma. Lactic acid is produced by Gram positive bacteria such as Lactobacillus and Pediococcus, which are present in dust and saliva. Acetic acid may be produced by several contaminants, including Acetobacter, Zymomonas, and yeast in the Kloeckera and Brettanomyces families. High levels of sour and acidic flavors generally indicate a sanitation problem, but they are an important part of the profile of the lambic, oud bruin and Berliner weiss styles, and to a lesser extent, Belgian white beers.
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